Kennedy, who was one of Albert Lehninger’s first graduate students. He was a postdoctoral fellow at the Harvard Medical School with Eugene P. Olaf College in 1964 and earned his PhD in Biochemistry at Stanford Medical School under Arthur Kornberg. Nelson, born in Fairmont, Minnesota, received his BS in Chemistry and Biology from St. White This page intentionally left blank About the Authors David L. FREEMAN AND COMPANY New York To Our Teachers Paul R. Cox Professor of Biochemistry University of Wisconsin–Madison W.H. Nelson Professor of Biochemistry University of Wisconsin–Madison Michael M. Freeman and Company 41 Madison Avenue New York, NY 10010 Houndmills, Basingstoke RG21 6XS, England LEHNINGER PRINCIPLES OF BIOCHEMISTRY F I F T H E D I T I O N David L. Freeman and Company All rights reserved Printed in the United States of America First printing W. Adam Steinberg using PDB ID 1I6H as modified by Seth Darst. RR DONNELLEY On the cover: RNA polymerase II from yeast, bound to DNA and in the act of transcribing it into RNA. This page intentionally left blank LEHNINGER PRINCIPLES OF BIOCHEMISTRY Publisher: Executive Editor: Senior Developmental Editor: Associate Director of Marketing: Marketing Director: Senior Media Editor: Managing Editor: Project Editor: Photo Editor: Photo Researcher: Text and Cover Designer: Page Makeup: Illustration Coordinator: Illustrations: Molecular Graphics: Production Coordinator: Composition: Manufacturing: SARA TENNEY KATHERINE AHR RANDI ROSSIGNOL DEBBIE CLARE JOHN BRITCH PATRICK SHRINER PHILIP McCAFFREY ELIZABETH GELLER BIANCA MOSCATELLI DENA BETZ VICKI TOMASELLI MARSHA COHEN SUSAN TIMMINS H.
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